Loma Linda University

Enrollment Information
Call us at: 909-558-1000

Faculty Directory
Gregory Nelson, PhD
Professor, Basic Sciences
School of Medicine
Professor, Radiation Medicine
School of Medicine
Publications    Scholarly Journals--Published
  • Jeffrey S. Willey • Shane A. J. Lloyd •(2011). Gregory A. Nelson • Ted A. Bateman  Ionizing Radiation and Bone Loss: Space Exploration and Clinical Therapy Applications. Clinic Rev Bone Miner Metab 9:54–62.  DOI 10.1007/s12018-011-9092-8 ( 8/2011 )
  • Sanchez MC, Nelson GA, Green LM. (2010) Effects of Protons and HZE Particles on Glutamate Transport in Astrocytes, Neurons and Mixed Cultures. Radiat Res. 174:669-78. ( 9/2010 )
  • Titova E, Adami A, Ostrowski R, Lalas S, Vlkolinsky R, Zhang JH, Nelson G, Obenaus A.  Radiation exposure prior to ischemia decreases lesion volume, brain edema and cell death. Acta Neurochir Suppl. 2010;106:51-3. PubMed PMID: 19812920 ( 9/2010 )
  • Xiao Wen Mao, XW,. Favre, CJ,  Fike, JR,  Kubinova, L, Anderson, E.  Campbell-Beachler, M, Jones, T,  Smith, A,  Rightnar, S  and GA Nelson (2010) High-LET Radiation-Induced Response of Microvessels in the Hippocampus.Rad. Res. 173: 486–493. ( 4/2010 )
  • Vlkolinsky R, Titova E, Krucker T, Chi BB, Staufenbiel M, Nelson GA, Obenaus A. (2010) Exposure to 56Fe-particle Radiation Accelerates Electrophysiological Alterations in the Hippocampus of APP23 Transgenic Mice. Radiat Res. 173:342-52. ( 3/2010 )
  • Med Phys. 2009 Aug;36(8):3714-23.  Dose response of CaF2:Tm to charged particles of different LET.  Moyers MF, Nelson GA.  PMID: 19746804 ( 8/2009 )
  • Eric R. Bandstra,ER, Thompson, RW, Nelson,GA, Willey, JS, Judex,S,  Cairns, M,  Benton, ER., Vazquez, ME, Carson, JA and TA Bateman (2009) Musculoskeletal Changes in Mice from 20–50 cGy of Simulated Galactic Cosmic Rays.  Rad. Res.172: 21–29. ( 7/2009 )
  • Vlkolinský, R, Krucker, T, Smith, A., Lamp, T, Nelson, G and Obenaus, A (2008)  Early effects of 56Fe radiation on normal and lipopolysaccharide-inhibited synaptic plasticity in mouse hippocampus. Radation Res. 169: 523 – 530. ( 9/2008 )
  • Lloyd, SAJ, Bandstra, ER, Travis, ND, NELSON, GA, Bourland, JD, Pecaut, MJ, Gridley, DS, Willey, JS and Bateman, TA. (2008) Spaceflight-relevant types of ionizing radiation and cortical bone: Potential LET effect? Adv. Space Res. 42: 1889 – 1897. ( 9/2008 )
  • Obenaus, A., Huang, L., Smith, A., Favre, CJ, NELSON, G and E Kendall (2008)  Magnetic Resonance Imaging and Spectroscopy of the Rat Hippocampus 1 Month after Exposure to 56Fe Particle Radiation.  Radiat. Res. 169(2): 149-161. ( 9/2008 )
  • Willey, JS,  Grilly, LS,  Howard, SH,  Pecaut, MJ.  Obenaus, A,  Gridley, DS,  NELSON, GA and T.A. Bateman. (2008) Bone Architectural and Structural Properties Following 56Fe26+ Radiation-Induced Changes in Body Mass. Radiat Res. 2008 170:201-207. ( 8/2008 )
  • Hintermueller, C, Coats, JS, Obenaus, AO, Nelson, G, Krucker, T and M Stampanoni (2008) 3D Quantification of Brain Microvessels Exposed to Heavy Particle Radiation.  Proc. 9th International Conference on X-ray Microscopy - XRM2008 ( 7/2008 )
  • Bandstra, E.,  Pecaut, M, Anderson, E, Willey, J, DeCarlo, F, Stock, S, Gridley, D, NELSON, GA,  Levine, H and T Bateman (2008)  Long-term Dose Response of Trabecular Bone in Mice to Proton Radiation.  Radiat. Res. 169: 607 – 614. ( 6/2008 )
  • Rola, R., Fishman, K, Baure, J, Rosi, S, Lamborn, K, Obenaus, A, NELSON, GA and JR Fike.  (2008)  Hippocampal Neurogenesis and Neuroinflammation after Cranial Irradiation with 56Fe Particles.  Radiat.  Res.  169: 626-632. ( 6/2008 )
  • Vlkolinský, R, Krucker, T, NELSON, GA and A Obenaus  (2008) 56Fe Radiation Reduces Neuronal Output and Attenuates Lipopolysaccharide-Induced Inhibition of Long-Term Potentiation in the Mouse Hippocampus.  Radiat. Res.  169: 523 – 530. ( 5/2008 )
  • Vlkolinsky, R, Krucker, T, Smith, A, Lamp, T, Nelson, GA, and A Obenaus . "Effects of Lipopolysaccharide on 56Fe Radiation-Induced Impairment of Synaptic Plasticity in Mouse Hippocampus." Radiation Research . (2007) 168: 462-470. ( 10/2007 )
  • Moyers, MF, Saganti, PB and GA Nelson . "EVA Space Suit proton and Electron Threshold Energy Measurements by XCT and Range Shifting." Radiation Measurements 41. (2007): 1216-1226. ( 1/2007 )
  • Coutrakon, GB, Benton, ER, Gridley, D, Hickey, T, Hubbard, J, Koss, P, Moyers, MF, NELSON, GA, Pecaut, M, Sanders, E and K Shahnazi . "Simulation of a 36 Hour Solar Particle Event at LLUMC Using a Proton Beam Scanning System. ." Nuclear Instruments and Methods. . (2007): -. ( 1/2007 )
  • Pecaut MJ, Dutta-Roy R, Smith AL, Jones TA, NELSON GA, Gridley DS.. "Acute effects of iron-particle radiation on immunity. Part I: Population distributions.." Radiat Res. 165. (2006): 68-77. ( 3/2006 )
    PMID: 16392964
  • Hamilton SA, Pecaut MJ, Gridley DS, Travis ND, Bandstra ER, Willey JS, NELSON GA, Bateman TA. .  A murine model for bone loss from therapeutic and space-relevant sources of radiation.  J Appl Physiol. 101. (2006): 789-793. PMID: 16741258 ( 3/2006 )
  • Gridley DS, Dutta-Roy R, Andres ML, NELSON GA, Pecaut MJ. . ") Acute effects of iron-particle radiation on immunity. Part II: Leukocyte activation, cytokines and adhesion." Radiat Res. 165. (2006): 78-87. ( 1/2006 )
    PMID: 16392965
  • Moyers, MF, Saganti, PB and GA NELSON . "EVA Space Suit Proton and Electron Threshold Energy Measurements by XCT and Range Shifting.." Int. J. Rad. Appl. Instrum. D. 41. (2006): 1216-1226. ( 1/2006 )
  • Rola R, Sarkissian V, Obenaus A, NELSON GA, Otsuka S, Limoli CL, Fike JR. "High-LET radiation induces inflammation and persistent changes in markers of hippocampal neurogenesis." Radiat Res. 164. (2005): 556-560. ( 10/2005 )
  • Green, L.M., Bianski, B.M., Murray, D.K., Rightnar, S.S. and G.A. NELSON . "Characterization of Accelerated Iron-Ion-Induced Damage in Gap Junction Competent and Incompetent Thyroid Follicular Cells." Radiat. Res. 163. (2005): 172-182. ( 10/2005 )
    PMID 15658893
  • Haerich P, NELSON GA, Pecaut MJ. . ") HZE radiation and dopaminergic modification of startle and prepulse inhibition in mice. ." Physiol Behav. 86. (2005): 103-110. ( 10/2005 )
    PMID: 16084538
  • Elmore, E., Lao, X.Y., Ko, M., Rigthnar, S., NELSON, G. and J.L. Redpath . "Neoplastic Transformation Induced by Low Dose Protons.." Int. J. Radiation Biology 81. (2005): 291-298. ( 4/2005 )
    The aim was to define the dose--response curve for high-energy proton-induced neoplastic transformation in vitro. The HeLa x skin fibroblast human hybrid cell assay was used to determine the frequency of neoplastic transformation following doses of 232 MeV protons (mean linear energy transfer, LET=0.44 keV microm(-1)) in the range 5-600 mGy. Proton irradiations were carried out at the Loma Linda University Proton Treatment Facility, CA, USA. The data indicate no evidence for induction of transformation below a dose of 100 mGy. At doses of 5 and 50 mGy, there is evidence for a possible suppression of transformation frequencies below that for spontaneous transformation. The shape of the dose--response curve for high-energy proton-induced transformation of the human hybrid cell line CGL1 does not follow a linear no-threshold model and shows evidence for a threshold as well as for possible suppression of transformation at doses
  • Green, L.M., Bianski, B.M., Murray, D.K., Rightnar, S.S. and G.A. NELSON. "Characterization of Accelerated Iron-Ion-Induced Damage in Gap Junction Competent and ?Incompetent Thyroid Follicular Cells." Radiat. Res. 163. (2005): 172-182. ( 1/2005 ) Link...
  • Rola, R., Otsuka, S., Obenaus, A., NELSON, G.A., Limoli, C.L., VandenBerg, S.R. and J.R. Fike . "Indicators of Hippocampal Neurogenesis are Altered by 56Fe Irradiation in a Dose Dependent Manner." Radiation Research 162. (2004): 442-446. ( 10/2004 )
  • Pecaut, M.J., Haerich, P., Zuccarelli Miller, C.N., Smith, A.L., Zendejas, E.D. and G.A. NELSON . "The Effects of Low-Dose, High-LET Radiation Exposure on Three Models of Behavior in C57Bl/6 Mice. ." Radiation Research 162. (2004): 148-156. ( 8/2004 )
    To investigate the behavioral consequences of exposure to whole-body irradiation such as might occur for astronauts during space flight, female C57BL/6 mice were exposed to 0, 0.1, 0.5 or 2 Gy accelerated iron ions (56Fe, Z = 26, beta = 0.9, LET = 148.2 keV/microm) of 1 GeV per nucleon using the Alternating Gradient Synchrotron at the Brookhaven National Laboratory. Animal testing began 2 weeks after exposure and continued for 8 weeks. Under these conditions, there were few significant effects of radiation on open-field, rotorod or acoustic startle activities at any of the times examined. The lack of radiation effects in these behavioral models appears to offer reassurance to NASA mission designers. These results suggest that there may be negligible effects of HZE radiation on many behaviors during a 2-8-week period immediately after radiation.
  • Pecaut MJ, Miller GM, Nelson GA, Gridley DS. Hypergravity-induced immunomodulation in a rodent model: hematological and lymphocyte function analyses. J Appl Physiol. 2004 Jul;97(1):29-38.. PubMed PMID: 14978009. ( 7/2004 )
  • Pecaut MJ, Nelson GA, Moyers MF, Rabin B, Gridley DS. "Out-of-field" effects  of head-localized proton irradiation on peripheral immune parameters. In Vivo. 2003 Nov-Dec;17(6):513-21. PubMed PMID: 14758715 ( 11/2003 - 12/2003 )
  • Nelson GA. Fundamental space radiobiology. Gravit Space Biol Bull. 2003 Jun;16(2):29-36. Review. PubMed PMID: 12959129. ( 6/2003 )
  • Pecaut MJ, Nelson GA, Peters LL, Kostenuik PJ, Bateman TA, Morony S, Stodieck LS, Lacey DL, Simske SJ, Gridley DS. Genetic models in applied physiology: selected contribution: effects of spaceflight on immunity in the C57BL/6 mouse. I. Immune population distributions. J Appl Physiol. 2003 May;94(5):2085-94. PubMed PMID: 12514166. ( 5/2003 )
  • Gridley DS, Nelson GA, Peters LL, Kostenuik PJ, Bateman TA, Morony S, Stodieck LS, Lacey DL, Simske SJ, Pecaut MJ. Genetic models in applied physiology: selected contribution: effects of spaceflight on immunity in the C57BL/6 mouse. II. Activation, cytokines, erythrocytes, and platelets. J Appl Physiol. 2003 May;94(5):2095-103. PubMed PMID: 12506046. ( 5/2003 )
  • Pecaut MJ, Gridley DS, Nelson GA. Long-term effects of low-dose proton radiation on immunity in mice: shielded vs. unshielded. Aviat Space Environ Med.  2003 Feb;74(2):115-24. PubMed PMID: 12602442. ( 2/2003 )
  • Higuchi Y, Nelson GA, Vazquez M, Laskowitz DT, Slater JM, Pearlstein RD. Apolipoprotein E expression and behavioral toxicity of high charge, high energy(HZE) particle radiation. J Radiat Res (Tokyo). 2002 Dec;43 Suppl:S219-24. PubMed PMID: 12793762. ( 12/2002 )
  • Green LM, Patel Z, Murray DK, Rightnar S, Burell CG, Gridley DS, Nelson GA. Cytoskeletal and functional changes in bioreactor assembled thyroid tissue organoids exposed to gamma radiation. J Radiat Res (Tokyo). 2002 Dec;43 Suppl:S213-8. PubMed PMID: 12793761. ( 12/2002 )
  • Nelson GA, Jones TA, Chesnut A, Smith AL. Radiation-induced gene expression in the nematode Caenorhabditis elegans. J Radiat Res (Tokyo). 2002 Dec;43 Suppl:S199-203. PubMed PMID: 12793758. ( 12/2002 )
  • Pecaut MJ, Haerich P, Zuccarelli CN, Smith AL, Zendejas ED, Nelson GA. Behavioral consequences of radiation exposure to simulated space radiation in the C57BL/6 mouse: open field, rotorod, and acoustic startle. Cogn Affect Behav Neurosci. 2002 Dec;2(4):329-40. PubMed PMID: 12641177. ( 12/2002 )
  • Gridley DS, Pecaut MJ, Green LM, Miller GM, Nelson GA. Hypergravity-induced immunomodulation in a rodent model: lymphocytes and lymphoid organs. J Gravit Physiol. 2002 Dec;9(2):15-27. PubMed PMID: 14638456. ( 12/2002 )
  • Green LM, Tran DT, Murray DK, Rightnar SS, Todd S, Nelson GA. Response of thyroid follicular cells to gamma irradiation compared to proton irradiation: II. The role of connexin 32. Radiat Res. 2002 Oct;158(4):475-85. PubMed PMID: 12236815. ( 10/2002 )
  • George K, Willingham V, Wu H, Gridley D, Nelson G, Cucinotta FA. Chromosome aberrations in human lymphocytes induced by 250 MeV protons: effects of dose, dose rate and shielding. Adv Space Res. 2002;30(4):891-9. PubMed PMID: 12539753. ( 9/2002 )
  • Gridley DS, Pecaut MJ, Nelson GA. Total-body irradiation with high-LET particles: acute and chronic effects on the immune system. Am J Physiol Regul Integr Comp Physiol. 2002 Mar;282(3):R677-88. PubMed PMID: 11832386. ( 3/2002 )
  • Pecaut MJ, Gridley DS, Smith AL, Nelson GA. Dose and dose rate effects of whole-body proton-irradiation on lymphocyte blastogenesis and hematological variables: part II. Immunol Lett. 2002 Jan 1;80(1):67-73. PubMed PMID: 11716967 ( 1/2002 )
  • Gridley DS, Pecaut MJ, Dutta-Roy R, Nelson GA. Dose and dose rate effects of  whole-body proton irradiation on leukocyte populations and lymphoid organs: part  I. Immunol Lett. 2002 Jan 1;80(1):55-66. PubMed PMID: 11716966. ( 1/2002 )
  • Green LM, Murray DK, Tran DT, Nelson GA, Shah MM, Luben RA. A spontaneously arising mutation in connexin32 with repeated passage of FRTL-5 cells coincides with increased growth rate and reduced thyroxine release. J Mol Endocrinol. 2001  Oct;27(2):145-63. PubMed PMID: 11564600. ( 10/2001 )
  • Nelson GA, Green LM, Gridley DS, Archambeau JO, Slater JM.  Research activities at the Loma Linda University and Proton Treatment Facility--an overview. Phys Med. 2001;17 Suppl 1:30-2.  PMID: 11770533 ( 9/2001 )
  • Pecaut MJ, Nelson GA, Gridley DS. Dose and dose rate effects of whole-body gamma-irradiation: I. Lymphocytes and lymphoid organs. In Vivo. 2001 May-Jun;15(3):195-208. PubMed PMID: 11491014. ( 5/2001 - 6/2001 )
  • Hartman PS, Hlavacek A, Wilde H, Lewicki D, Schubert W, Kern RG, Kazarians GA, Benton EV, Benton ER, Nelson GA. A comparison of mutations induced by accelerated iron particles versus those induced by low earth orbit space radiation in the FEM-3 gene of Caenorhabditis elegans. Mutat Res. 2001 Mar 1;474(1-2):47-55. PubMed PMID: 11239962. ( 3/2001 )
  • Green LM, Murray DK, Bant AM, Kazarians G, Moyers MF, Nelson GA, Tran DT. Response of thyroid follicular cells to gamma irradiation compared to proton irradiation. I. Initial characterization of DNA damage, micronucleus formation, apoptosis, cell survival, and cell cycle phase redistribution. Radiat Res. 2001 Jan;155(1 Pt 1):32-42. PubMed PMID: 11121213. ( 1/2001 )
  • Kajioka EH, Andres ML, Mao XW, Moyers MF, Nelson GA, Gridley DS. Hematological and TGF-beta variations after whole-body proton irradiation. In Vivo. 2000 Nov-Dec;14(6):703-8. PubMed PMID: 11204485. ( 11/2000 - 12/2000 )
  • Gridley DS, Pecaut MJ, Miller GM, Moyers MF, Nelson GA. Dose and dose rate effects of whole-body gamma-irradiation: II. Hematological variables and cytokines. In Vivo. 2001 May-Jun;15(3):209-16. PubMed PMID: 11491015. ( 5/2001 - 6/2000 )
  • Kajioka EH, Andres ML, Nelson GA, Gridley DS. Immunologic variables in male and female C57BL/6 mice from two sources. Comp Med. 2000 Jun;50(3):288-91. PubMed PMID: 10894494. ( 6/2000 )
  • Kajioka EH, Andres ML, Li J, Mao XW, Moyers MF, Nelson GA, Slater JM, Gridley DS. Acute effects of whole-body proton irradiation on the immune system of the mouse. Radiat Res. 2000 May;153(5 Pt 1):587-94. PubMed PMID: 10790280. ( 5/2000 )
  • Kajioka EH, Gheorghe C, Andres ML, Abell GA, Folz-Holbeck J, Slater JM, Nelson GA, Gridley DS. Effects of proton and gamma radiation on lymphocyte populations and acute response to antigen. In Vivo. 1999 Nov-Dec;13(6):525-33. PubMed PMID: 10757049. ( 11/1999 - 12/1999 )
  • Li J, Andres ML, Fodor I, Nelson GA, Gridley DS. Evaluation of pGL1-TNF-alpha therapy in combination with radiation. Oncol Res. 1998;10(7):379-87. PubMed PMID: 10063972. ( 11/1998 - 12/1998 )
  • NELSON, G.A., Kazarians, G.A. & Schubert, W.W.  (1997)  Heavy Ion Effects on Caenorhabditis elegans.  SAE Technical Paper Series 972373. Society of Automotive Engineers, Inc. Warrendale, PA. ( 9/1997 )
  • NELSON, G.A., Schubert, W.W., Kazarians, G.A., Richards, G.F., Benton, E.V., Benton, E.R.& Henke, R. (1994) Radiation Effects in Nematodes:  Results from IML-1 Experiments.  Advances in Space Research 14 (10): 87‑91. ( 9/1994 )
  • NELSON, G.A., Schubert,W.W., Kazarians, G.A. & Richards, G.F.  (1994) Development and Chromosome Mechanics in Nematodes:  Results from IML-1.  Advances in Space Research  14(8): 209-214. ( 9/1994 )
  • NELSON, G.A., Schubert, W.W., & Marshall, T.M.  (1992) Radiobiology studies with the nematode Caenorhabditis elegans.  Genetic and Developmental Effects of High LET Radiation.  Nuclear Tracks and Radiation Measurements  20: 277-232. ( 9/1992 )
  • Johnson, T.E., & NELSON, G.A.  (1991) Caenorhabditis elegans: A Model System for Space Biology Studies.  Experimental Gerontology  26:299-309. ( 9/1991 )
  • NELSON, G.A. (1990) Radiation Biology of HZE Particles. Proceedings of the 41st Congress of the International Astronautical Federation. October 6-12, 1990, Dresden, GDR. Paper IAA-90-548, International Astronautical Federation, Paris. ( 9/1990 )
  • Petersen, G.R., W.W. Schubert, G.F. Richards and G.A. NELSON (1990)  Yeasts Producing Exopolysaccharides with Drag-Reducing Activity.  Enzyme Microb. Technol: 12: 255 -259. ( 9/1990 )
  • Petersen, G.R., G.A. NELSON, C.A. Cathey, and G.G. Fuller (1989) Rheologically Interesting Polysaccharides from Yeasts.  Applied Biochemistry and Biotechnology. 20/21: 845-867. ( 9/1989 )
  • NELSON, G.A., Schubert, W.W., Marshall, T.M., Benton, E.R. & Benton, E.V.  (1989) Radiation effects in Caenorhabditis elegans, mutagenesis by high and low LET ionizing radiationMutation Research 212:181-192. ( 9/1989 )
  • Coohill, T.P., Marshal, T.M.,  Schubert, W.W., & NELSON, G.A.  (1988) Ultraviolet mutagenesis of radiation-sensitive (rad) mutants of the nematode C. elegans.  Mutation Research  209: 99-106. ( 9/1988 )
  • NELSON, G.A., Andrews, M.L. & Karnovsky, M.J.  (1983) Control of Erythrocyte Shape by Calmodulin.  Journal of Cell Biology  96: 730‑735. ( 9/1983 )
  • NELSON, G.A., Andrews, M.L. and Karnovsky, M.J.  (1982) The Participation of Calmodulin in Immunoglobulin Capping.  Journal of Cell Biology  95: 771‑780. ( 9/1982 )
  • Ward, S., T.M. Roberts, G.A. NELSON and Y. Argon (1982) The Development and Motility of C.elegans Spermatozoa. J. Nematology 14: 259-266. ( 9/1982 )
  • Ward, S., NELSON, G.A., & Argon, Y.  (1981) Sperm morphogenesis in wild type and fertilization defective mutants of C. elegans.  Journal of Cell Biology  91: 26‑44. ( 9/1981 )
  • Ward, S., G.A. NELSON and Y. Argon (1981) Sperm Morphogeneiss in Wild Type and Fertilization Defective Mutants of C. elegans.  Journal of Cell Biology 91: 26-44. ( 9/1981 )
  • NELSON, G.A. and S. Ward (1981) Amoeboid Motility and Actin in Ascaris lumbricoides Sperm.  Experimental Cell Research 131:  149-160. ( 9/1981 )
  • NELSON, G.A., & Ward, S.  (1980) Vesicle fusion, pseudopod extension and amoeboid motility are induced in nematode spermatids by the ionophore monensin.  Cell 19: 457‑464. ( 9/1980 )
  • NELSON, G.A. and S. Ward (1980) Vesicle Fusion, Pseudopod Extension and Amoeboid Motility are Induced in Nematode Spermatids by the Ionophore Monensin.  Cell 19:  457-464. ( 9/1980 )
  • NELSON, G.A., K.K. Lew and S. Ward (1978) Intersex, A Temperature Sensitive Mutant of the Nematode Caenorhabditis elegans.  Developmental Biology 66:  386-409. ( 9/1978 )
  • G. A. NELSON, K. K. Lew, and S. Ward  (1978) Intersex, a Temperature-Sensitive Mutant of the Nematode  Caenorhabditis elegans.  Developmental Biology.  66: pp. 386-409. ( 9/1978 )
  • NELSON, G.A., and J.P. Revel (1975) Scanning Electron Microscopic Study of Cell Movements in the Corneal Endothelium of the Avian Embryo.  Developmental Biology 42:  315-33. ( 9/1975 )
  Scholarly Journals--Accepted
  • Jeffrey S. Willey, Shane A.J. Lloyd, Gregory A. Nelson, Ted A. Bateman. (2011)  Space Radiation and Bone Loss.  Gravit Space Biol Bull.  in press. ( 12/2011 )
  Books and Chapters
  • Gridley DS, Green LM, NELSON GA, Pecaut MJ & Slater JM. . Cancer - Radiation-Induced Damage. . Therapeutic Applications of Superoxide Dismutase and its Mimetics. : Landers Bioscience-Riley , 2005. ( 1/2005 )
  • NELSON, G.A., Schubert, W.W., Kazarians, G.A., Kern, R.G., Schranck, D., Hartman, P.S., Hlavacek, A., Benton, E.V. & Benton, E.R.(1999)  Modification of Radiogenic Damage by Microgravity, In. Biorack on Shuttle to Mir. European Space Agency Special Publication:  ESA-SP-1222. ESA, Noordwijk, Netherlands. ( 9/1999 )
  • Hartman, P. & NELSON, G.A.  (1997) Processing of DNA Damage in the Nematode Caenorhabditis elegans.  Chapter 24.  In: DNA Damage and Repair.  Biochemistry, Genetics, and Cell Biology.  Eds.,  Hoekstra, M.F.& Nickoloff, J.A.. Humana Press, Inc.,  Totowa, N.J.:  pp. 557-576. ( 9/1997 )
  • NELSON, G. A.  (1995) Radiation in Microgravity.  Chapter 34.  In: Handbook of Physiology.  Sec. 4.  Adaptation to the Environment.  Fregly, M.J. & Blatteis, C.M., Editors. pp. 785-798.  Oxford University Press, New York. ( 9/1995 )
  • NELSON, G.A., W.W. Schubert, Kazarians,G.A., Richards, G.F., Benton, E.V. Benton, E.R. & Henke, R. (1995) Genetic and Molecular Dosimetry of HZE Radiation.  In: Biorack on Spacelab IML-1.  European Space Agency Publication ESA SP-1162.  European Space Agency, Noordwijk, The Netherlands.  pp 41-50.  [ISBN 92-9092-085-8]. ( 9/1995 )
  • NELSON, G.A., Schubert, W.W.,& Kazarians, G.A.  (1992) Heavy ion radiobiology of the nematode Caenorhabditis elegans.  Radiation Research A Twentieth Century Perspective.  Dewey, W., Edinton, M., Fry, R., Hall, E. and Whitmore, G. eds.  Academic Press, Inc., San Diego, pp. 493‑497. ( 9/1992 )
  • NELSON, G., T. Marshall and W. Schubert (1989) Genetic and Developmental Effects of High LET Radiation on the Nematode C. elegans. Proceedings of the 27th Hanford Life Sciences Symposium. Multilevel Health Effects Research: From Molecules to Man. Ed. J.F. Park and R.A. Pelroy. Richland, Washington. October 18 - 21, 1988. Battelle Press, Columbus, Ohio. ( 9/1988 )
  Non-Scholarly Journals
  • Ward, S., G.A. NELSON and Y. Argon (1980).  Genetic Specification of Cellular Morphology.  in Carnegie Institution Yearbook 79, 1979-1980.  Carnegie Institution of Washington, Washington, DC pp. 37-41. ( 9/1980 )
  • NELSON, G.A. (1979) Motility of Nematode Sperm.  in Carnegie Institution Yearbook 78, 1978-1979.  Carnegie Insitution of Washington, Washington, D.C. pp. 62-66. ( 9/1979 )
  • NELSON, G.A., K.K. Lew and S. Ward (1978) Temperature Sensitive Intersex Mutant in Carnegie Institution Yearbook 77, 1977-1978.  Carnegie Institution of Washington.  Washington, D.C., pp. 58-61. ( 9/1978 )